Whilst there is a membrane, it generally doesn't form a complete seal.
So, from personal experience, the draughts find their way in around the door handle, the window winders, past the interior window rubbers, and anywhere else it has any chance. Sealing the trailing edge door vents with duct tape siginficantly reduces this.
The problem with motorhome conversions is that any fixed ventilation (of which there is much on many models) in the habitation area behind the cab, combined with the body shape, tends to alter the cab ventilation characteristics dramatically.
Gaps in the cab area which would normally allow air OUT in a panel van, tend instead to suck air IN and pass it back, to be ventilated further back.
I can assure you that in two vans of similar vintage, as I've already said, the biggest problem I had was around the area of the inertia reel for the seat belt - with a block of foam rubber inserted in the box section below and behind making a huge difference. There is a complete void through to the underneath of the van!
(Incidentally, I have noticed the same characteristic in the current Transit).